1. Department of Science; Faculty of Management, Science &Technology; Open Universiteit; 6401 DL Heerlen The Netherlands
2. Netherlands Centre for Biodiversity Naturalis; 2300 RA Leiden The Netherlands
3. Department of Malacology; Academy of Natural Sciences; Philadelphia PA 19130 USA
4. USDA-APHIS-PPQ; Department of Malacology; Academy of Natural Sciences; Philadelphia PA 19130 USA