1. College of Veterinary Medicine South China Agricultural University Guangzhou China
2. Key Laboratory of Animal Vaccine Development Ministry of Agriculture Guangzhou China
3. National and Regional Joint Engineering Laboratory for Medicament of Zoonosis Prevention and Control Guangzhou China
4. Key Laboratory of Zoonosis Prevention and Control of Guangdong Province Guangzhou China
5. Shanghai Veterinary Research Institute (SHVRI)Chinese Academy of Agricultural Sciences (CAAS) Shanghai China