1. Key Laboratory of Zoonotic of Liaoning Province College of Animal Science and Veterinary Medicine Shenyang Agricultural University Shenyang Liaoning Province P. R. China
2. Institute of Geographic Sciences and Natural Resources Research Chinese Academy of Sciences Beijing P. R. China
3. Plague and Brucellosis Prevention and Control Base Chinese Center for Disease Control and Prevention Baicheng P. R. China
4. School of Public Health Sun Yat‐sen University Guangzhou P. R. China