1. Key Laboratory of Aquaculture Disease Control; Ministry of Agriculture, and State Key Laboratory of Freshwater Ecology and Biotechnology; Institute of Hydrobiology; Chinese Academy of Sciences; Wuhan 430072 China
2. Departamento de Microbiología y Parasitología; Facultad de Farmacia; Universidad Complutense de Madrid; Plaza Ramóny Cajal s/n 28040 Madrid Spain
3. Department of Biological Sciences; Northern Arizona University; Flagstaff Arizona 86011