1. Japan Sea National Fisheries Research Institute, Japan Fisheries Research and Education Agency Niigata Japan
2. Graduate School of BioresourcesMie University Tsu City Japan
3. Minamisanriku‐cho Government Office Minamisanriku‐cho Japan
4. Graduate School of Agricultural ScienceTohoku University Sendai Japan
5. Department of Ocean Sciences, Tokyo University of Marine Science and Technology Tokyo Japan