1. Institut des Sciences de l’Évolution, UMR 5554, CNRS; Université Montpellier 2; Montpellier cedex 5 Place Eugène Bataillon France 34095
2. INRA, CBGP (UMR INRA - IRD - IRAD - Montpellier SupAgro); Campus international de Baillarguet, CS 30016, F-34988 Montferrier-sur-Lez Cedex; France
3. Department of Biological Sciences; The University of Tokyo; Hongo 7-3-1 Bunkyoku Tokyo 113-0033 Japan
4. CEFE, CNRS UMR 5175, 1919 route de Mende; F-34293 Montpellier cedex 5; France