1. Island Ecology and Evolution Research Group; Instituto de Productos Naturales y Agrobiología (IPNA-CSIC); C/Astrofísico Francisco Sánchez 3 La Laguna Tenerife, Canary Islands 38206 Spain
2. Biology Department; City College of New York; New York NY 10031 USA
3. The Graduate Center; City University of New York; New York NY 10016 USA