1. Bone & Joint Research Unit; William Harvey Research Institute; Barts and The London School of Medicine and Dentistry; Queen Mary University of London; London; UK
2. Division of Cancer Studies; School of Medicine; King's College London; London; UK
3. Immunology Laboratory; University of Brest Medical School; Brest; France
4. Centre for Rheumatology; UCL Division of Medicine; University College London; London; UK