1. Department of Biology; Penn State University; University Park PA 16802 USA
2. School of Ocean and Earth Science and Technology; University of Hawai‘i at Mānoa; HI 96822 USA
3. Institute of Marine Sciences; University of Dar es Salaam; Zanzibar Tanzania
4. Biodiversity Research Center/Taiwan International Graduate Program (TIGP)-Biodiversity; Academia Sinica; Nangang Taipei Taiwan
5. Red Sea Research Center; King Abdullah University of Science and Technology (KAUST); Thuwal Saudi Arabia