1. School of Plant Biology, Faculty of Science; University of Western Australia; Crawley Western Australia 6009 Australia
2. UWA Oceans Institute; University of Western Australia; Crawley Western Australia 6009 Australia
3. Virginia Institute of Marine Science; College of William and Mary; Gloucester Point VA 23062 U.S.A.
4. School of Earth and Environment; University of Western Australia; Crawley Western Australia 6009 Australia
5. ARC Centre of Excellence for Coral Reef Studies; James Cook University Townsville; Queensland 4811 Australia
6. Kings Park and Botanic Garden; West Perth Western Australia 6005 Australia