1. Department of Fisheries, Faculty of Marine Natural Resources Khorramshahr University of Marine Science and Technology Khorramshahr Iran
2. Department of Zoology University of Otago Dunedin New Zealand
3. Department of Natural Resources Isfahan University of Technology Isfahan Iran
4. Institute of Biotechnology and Bioengineering Isfahan University of Technology Isfahan Iran
5. Fisheries Department, Faculty of Natural Resources University of Guilan Guilan Iran