1. School of Biology; Newcastle University; Newcastle upon Tyne NE1 7RU UK
2. IUCN SSC Conservation Breeding Specialist Group; 12101 Johnny Cake Ridge Road Apple Valley MN 55124 USA
3. IUCN SSC Conservation Breeding Specialist Group; European Office, Copenhagen Zoo, p/a Annuntiatenstraat 6 2170 Merksem Belgium
4. European Association of Zoos and Aquaria; PO Box 20164 1000 HD Amsterdam The Netherlands