1. Percy FitzPatrick Institute, DST/NRF Centre of ExcellenceUniversity of Cape Town Rondebosch Cape Town 7701 South Africa
2. Institute of ZoologyZoological Society of London Regent's Park London NW1 4RY UK
3. Centre for Biodiversity and Environment Research, Department of Genetics, Evolution and EnvironmentUniversity College London London WC1E 6BT UK
4. School of Geography and the EnvironmentUniversity of Oxford South Parks Road Oxford OX1 3PY UK
5. South African National ParksScientific Services Private Bag X1021 Phalaborwa 1390 South Africa
6. ARC Centre of Excellence for Coral Reef StudiesJames Cook University Townsville QLD 4811 Australia