1. Department of Fish and Wildlife SciencesUniversity of Idaho Moscow ID 83844 USA
2. Department of Biological Sciences University of Alberta Edmonton AB T6G0C5 Canada
3. Department of Biology and WildlifeUniversity of Alaska Fairbanks Fairbanks AK 99775 USA
4. UAF School of ManagementUniversity of Alaska Fairbanks Fairbanks AK 99775 USA
5. School of Environmental and Forest SciencesUniversity of Washington WA 98195 USA