1. Departament de Biologia Marina i Oceanografia Institut de Ciències del Mar (CSIC) Pg. Marítim de la Barceloneta, 37‐49 Barcelona Catalonia08003Spain
2. Living Systems Institute University of Exeter Stocker Road ExeterEX4 4QDUK
3. Department of Zoology University of Oxford 11a Mansfield Road OxfordOX1 3SZUK