1. Department of Wildlife, Fisheries, and Aquaculture Mississippi State University Mississippi State Mississippi USA
2. Laboratoire Evolution et Diversité Biologique (EDB), UMR5174, CNRS, Institut de Recherche pour le Développement (IRD) Université Toulouse 3 Paul Sabatier Toulouse France
3. Museum of Paleontology University of Michigan Ann Arbor Michigan USA
4. Department of Ecology and Evolutionary Biology University of Michigan Ann Arbor Michigan USA