1. Institute of Life Course and Medical Sciences University of Liverpool Liverpool UK
2. The Medical Research Council/Versus Arthritis Centre for Integrated Research into Musculoskeletal Aging CIMA University of Liverpool Liverpool UK
3. Discipline of Physiology School of Medicine National University of Ireland Galway Ireland
4. Department of Oncology and Metabolism Healthy Lifespan Institute and the Centre for Integrated Research in Musculoskeletal Aging University of Sheffield Sheffield UK