1. School of Biological Sciences University of Bristol Bristol BS8 1TQUK
2. Department of Life Sciences Imperial College London Silwood Park CampusAscot SL5 7PYUK
3. EMBL/CRG Systems Biology Research Unit, Centre for Genomic Regulation (CRG) The Barcelona Institute of Science and Technology Dr. Aiguader 88Barcelona 08003Spain
4. Universitat Pompeu Fabra (UPF) 08002Barcelona Spain
5. Centre for Biodiversity and Environmental Research, Department of Genetics, Evolution and Environment University College London London WC1E 6BTUK