1. Princeton School of Public and International Affairs Princeton University Princeton NJ USA
2. Key Laboratory of Animal Ecology and Conservation Biology Institute of Zoology Chinese Academy of Sciences Beijing China
3. Key Laboratory of Adaptation and Evolution of Plateau Biota Northwest Institute of Plateau Biology Chinese Academy of Sciences Xining China
4. Key Laboratory of Zoological Systematics and Evolution Institute of Zoology Chinese Academy of Sciences Beijing China
5. Department of Ecology and Evolutionary Biology Princeton University Princeton NJ USA