1. Ecology of Terrestrial and Aquatic Systems Laboratory (EcoSTAq), Department of Biology; Badji Mokhtar University; Annaba Algeria
2. Université Paul-Valéry Montpellier UMR 5175 CEFE (Centre d’Ecologie fonctionnelle et évolutive), Route de Mende; F-34199 Montpellier Cedex France
3. Department of Biology; Jordan University of Science and Technology; Irbid Jordan