1. Museum of Comparative Zoology Department of Organismic and Evolutionary Biology Harvard University 26 Oxford Street Cambridge MA 02138 USA
2. Department of Evolutionary Biology, Ecology, and Environmental Sciences Faculty of Biology, and Biodiversity Research Institute (IRBio) University of Barcelona 643 Diagonal Av. Barcelona, Catalonia 08028 Spain
3. Phylogenetic Systematics and Evolution Research Group Section of Taxonomy and Evolution Department of Natural History University Museum of Bergen University of Bergen Bergen PB7800 5020 Norway