1. Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, China
2. Graduate University of Chinese Academy of Sciences, Beijing, 100049, China
3. State Key Laboratory of Genetic Resources and Evolution, Kunming Institute of Zoology, Chinese Academy of Sciences, Kunming 650223, China
4. Centre for Biodiversity and Conservation Biology, Royal Ontario Museum, 100 Queen’s Park, Toronto, Ontario M5S 2C6, Canada