1. Animal Breeding and Genomics Centre, Wageningen University, De Elst 1, Wageningen, 6708 WD, the Netherlands
2. Department of Animal Ecology, Netherlands Institute of Ecology (NIOO‐KNAW), PO Box 50, 6700 AB Wageningen, the Netherlands
3. Department of Animal and Plant Sciences, University of Sheffield, Sheffield S10 2TN, UK
4. Edward Grey Institute, Department of Zoology, University of Oxford, Oxford OX1 3PS, UK