1. Department of Genetics, Evolution and Environment, University College London, 4 Stephenson Way, London NW1 2HE, UK
2. NERC Centre for Population Biology, Imperial College London, Silwood Park Campus, Ascot, Berkshire SL5 7PY, UK; CNRS, UMR 7205, Muséum National d'Histoire Naturelle, 16 Rue Buffon, CP39, 75005 Paris, France
3. FAS Center for Systems Biology, Harvard University, 52 Oxford St, Northwest Lab Room 458.40‐3, Cambridge, MA 02138, USA
4. Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK
5. Wissenschaftskolleg zu Berlin, Wallotstraße 19, 14193 Berlin, Germany