1. Department of Zoology University of Cambridge Cambridge CB2 3EJ United Kingdom
2. Centre for Ecology and Conservation, College of Life and Environmental Sciences University of Exeter Penryn TR10 9FE United Kingdom
3. FitzPatrick Institute of African Ornithology, DST‐NRF Centre of Excellence, Department of Biological Sciences University of Cape Town Cape Town 7701 South Africa
4. Current Address: Cambridge Institute of Therapeutic Immunology & Infectious Disease (CITIID), Jeffrey Cheah Biomedical Centre, Cambridge Biomedical Campus University of Cambridge Cambridge CB2 0AW United Kingdom