1. College of Fisheries; Guangdong Ocean University; Zhanjiang China
2. Guangdong Provincial Key Laboratory of Pathogenic Biology and Epidemiology for Aquatic Economic Animals; Zhanjiang China
3. Guangdong Key Laboratory of Control for Diseases of Aquatic Economic Animals; Zhanjiang China
4. Department of Fisheries and Aquatic Resources Management; University for Development Studies; Tamale Ghana
5. Langye Animal Husbandry Company Limited; Gaozhou City Guangdong province