1. Academic Unit of Surgery, School of Medicine, Dentistry and Nursing University of Glasgow Glasgow UK
2. Department of Surgery Raigmore Hospital Inverness UK
3. Centre for Research into Cancer Prevention and Screening, School of Medicine, Ninewells, Hospital and Medical School University of Dundee Dundee UK
4. Department of Blood Sciences, Ninewells Hospital and Medical School Dundee UK
5. Population Health & Genomics, School of Medicine, Ninewells Hospital and Medical School University of Dundee Dundee UK