1. Department of Botany; Institute of Ecology and Earth Sciences; University of Tartu; Lai 40 Tartu EE-51005 Estonia
2. Department of Biology; University of British Columbia - Okanagan Campus; Kelowna British Columbia V1V 1V Canada
3. Evolution & Ecology Research Centre; School of Biological; Earth and Environmental Sciences; University of New South Wales; Sydney NSW 2052 Australia