1. Centre for Biodiversity Dynamics; Department of Biology; Norwegian University of Science and Technology, NTNU; 7491 Trondheim Norway
2. School of Biological Sciences; University of Portsmouth; King Henry Building, King Henry I Street Portsmouth PO1 2DY UK
3. Department of Biology; Norwegian University of Science and Technology, NTNU; 7491 Trondheim Norway
4. Institute of Arctic Biology; University of Alaska; Fairbanks AK 99775 USA
5. School of Biology and Center of Excellence in Phylogeny of Living Organisms; University of Tehran; 14155-6455 Tehran Iran