1. Department of Biological Sciences; University of Cape Town; P/Bag X3, Rondebosch Cape Town 7701 South Africa
2. Institute of Evolutionary Biology and Environmental Sciences; University of Zurich; Winterthurerstrasse 190 Zurich CH-8057 Switzerland
3. School of Life Sciences; University of KwaZulu-Natal; P/Bag X01, Scottsville Pietermaritzburg 3209 South Africa