1. Environmental and Marine Biology Åbo Akademi University Artillerigatan 6 20520 Åbo Finland
2. Finnish Environmental Institute/Marine Research Centre PB 140 00251 Helsinki Finland
3. Department of Biological and Environmental Sciences University of Gothenburg PB 461 SE 40530 Göteborg Sweden
4. Department of Biological and Environmental Sciences – Tjärnö University of Gothenburg SE 45296 Strömstad Sweden