1. Centre for Environmental Studies Department of Geography, Geoinformatics and Meteorology University of Pretoria Pretoria South Africa
2. CIRAD UMR Animal Santé Risque et Ecosystèmes (ASTRE) Montpellier France
3. ASTRE CIRAD INRA University of Montpellier Montpellier France
4. Department of Zoology and Entomology Mammal Research Institute University of Pretoria Pretoria South Africa
5. Department of Production Animal Studies Faculty of Veterinary Sciences University of Pretoria Pretoria South Africa