1. Laboratory of Animal Breeding and Genetics; Graduate School of Agriculture; Kyoto University; Kyoto Japan
2. Department of Agrobiology; Faculty of Agriculture; Niigata Japan
3. Field Center for Sustainable Agriculture and Forestry; Niigata University; Niigata Japan
4. Sado Japanese Crested Ibis Conservation Center; Sado Japan
5. Yamashina Institute for Ornithology; Chiba Japan