1. Ecological Genetics Research Unit, Organismal and Evolutionary Biology Research Programme, Faculty of Biological and Environmental Sciences University of Helsinki Helsinki 00014 Finland
2. Department of Organismic and Evolutionary Biology and Museum of Comparative Zoology Harvard University Cambridge Massachusetts 02138 USA
3. Department of Biochemistry, Genetics, and Immunology Universidade de Vigo Vigo 36310 Spain
4. Lammi Biological Station University of Helsinki Lammi 16900 Finland
5. Kilpisjärvi Biological Station University of Helsinki Kilpisjärvi 99490 Finland
6. Area of Ecology and Biodiversity, School of Biological Sciences The University of Hong Kong Pok Fu Lam Hong Kong