1. US Geological Survey National Wetlands Research Center Lafayette LA 70506 USA
2. School of Biological Sciences University of Queensland St Lucia Qld 4072 Australia
3. US Geological Survey Patuxent Wildlife Research Center Beltsville MD 20705 USA
4. Office of Environment and Heritage NSW Department of Premier and Cabinet Sydney NSW 1232 Australia
5. Key Laboratory of Ministry of Education for Coastal and Wetland Ecosystems College of the Environment and Ecology Xiamen University Xiamen 361005 Fujian China