1. Organisms and Environment Research Division; Cardiff School of Biosciences; Cardiff University; Sir Martin Evans Building, Museum Avenue Cardiff CF10 3AX Wales UK
2. Centre d'Ecologie Fonctionnelle et Evolutive CEFE; UMR 5175; CNRS; Université de Montpellier; Université Paul-Valéry Montpellier, EPHE; 1919 route de Mende 34293 Montpellier Cedex 5 France