1. Frontier Research Institute for Interdisciplinary Sciences; Tohoku University; 6-3 Aoba Aramaki Aoba Sendai Miyagi 980-8578 Japan
2. Division of Ecology and Evolutionary Biology; Graduate School of Life Sciences; Tohoku University; 6-3 Aoba Aramaki Aoba Sendai Miyagi 980-8578 Japan
3. Field Science Center; Graduate School of Agricultural Science; Tohoku University; 232-3 Yomogida Naruko-onsen Osaki Miyagi 989-6711 Japan
4. Department of Cell Proliferation; United Center for Advanced Research and Translational Medicine; Graduate School of Medicine; Tohoku University; 2-1 Seiryo Aoba Sendai 980-8575 Japan