1. Institute of Life Science Swansea University Medical School Swansea UK
2. MRC Integrative Epidemiology Unit University of Bristol Oakfield House Bristol UK
3. Cellular and Molecular Medicine University of Bristol Biomedical Sciences Building Bristol UK
4. Alberta Respiratory Centre (ARC) Department of Medicine University of Alberta Edmonton AB Canada
5. Division of Cancer and Genetics Department of Haematology School of Medicine Cardiff University Cardiff UK