1. Institute of Evolution and Marine Biodiversity; Ocean University of China; Qingdao 266003 China
2. Key Laboratory of Aquatic Biodiversity and Conservation of Chinese Academy of Sciences; Institute of Hydrobiology; Chinese Academy of Sciences; Wuhan 430072 China
3. Department of Biological Sciences; Boise State University; Boise 83725 ID USA
4. School of Life Sciences; East China Normal University; Shanghai 200062 China
5. Laboratory for Marine Biology and Biotechnology; Qingdao National Laboratory for Marine Science and Technology; Qingdao 266237 China