1. Department of Natural Sciences; Unitec Institute of Technology; Auckland 1025 New Zealand
2. Institute of Fundamental Sciences; Massey University; Palmerston North 4474 New Zealand
3. UMR 6553 ECOBIO; Université de Rennes 1; CNRS; F-35042 Rennes CEDEX France
4. Alterra; Wageningen UR; 6700AA Wageningen The Netherlands
5. Allan Herbarium; Landcare Research; Lincoln 7640 New Zealand