1. Department of Ecology and Evolutionary Biology University of Tennessee Knoxville Tennessee USA
2. Illinois Natural History Survey, Prairie Research Institute University of Illinois Champaign USA
3. Institute of Oceanography National Taiwan University Taipei Taiwan
4. Ushimado Marine Institute, Okayama University 130‐17 Kashino, Setouchi Okayama Japan
5. Key Laboratory of Animal Ecology and Conservation Biology Chinese Academy of Sciences Beijing China