1. UMR7205 Institut de Systématique; Evolution, Biodiversité CNRS MNHN UPMC EPHE; Sorbonne Universités; Muséum National d'Histoire Naturelle; CP 51, 57 rue Cuvier F-75231 Paris Cedex 05 France
2. North Carolina Museum of Natural Sciences; Raleigh NC USA
3. Vertebrates Section; Museums Victoria Melbourne VIC 3001 Australia
4. 6 Stratton Road Saltford Bristol BS31 3BS UK
5. Museum of Vertebrate Zoology and Department of Integrative Biology; 3101 Valley Life Sciences Building; University of California; Berkeley CA 94720-3160 USA