1. Institute of Oceanology; Chinese Academy of Sciences; Qingdao China
2. Center for Ocean Mega-Science; Chinese Academy of Sciences; Qingdao China
3. University of Chinese Academy of Sciences; Beijing China
4. Institute of Marine Biology and Center of Excellence for the Oceans; National Taiwan Ocean University; Keelung Taiwan
5. Laboratory for Marine Biology and Biotechnology; Qingdao National Laboratory for Marine Science and Technology; Qingdao China