1. Research School of Biology The Australian National University RN Robertson Building, 46 Sullivans Creek Road Acton (Canberra) ACT 2601 Australia
2. School of Biological Sciences The University of Queensland Brisbane Qld 4072 Australia
3. School of Natural Sciences The University of Tasmania Private Bag 55 Hobart Tas 7001 Australia
4. Graduate School of Agriculture Kyoto University Kyoto 606‐8502 Japan
5. Graduate School of Human and Environmental Studies Kyoto University Kyoto 606‐8501 Japan