1. State Key Laboratory of Soil and Sustainable Agriculture Institute of Soil Science Chinese Academy of Sciences Nanjing 210008 China
2. State Key Laboratory of Crop Genetics and Germplasm Enhancement Cytogenetics Institute Nanjing Agricultural University/Jiangsu Collaborative Innovation Center for Modern Crop Production Nanjing Jiangsu 210095 China
3. Institute of Food Crops Jiangsu Academy of Agricultural Sciences Nanjing Jiangsu 210014 China
4. School of BioSciences The University of Melbourne Parkville Vic. 3010 Australia
5. Faculty of Land and Food Systems University of British Columbia Vancouver BC V6T 1Z4 Canada
6. National Centre for Plant Gene Research (Beijing) Institute of Genetics and Developmental Biology Chinese Academy of Sciences Beijing 100049 China