1. Department of Ecology and Evolutionary Biology University of Connecticut Storrs Connecticut 06269
2. Current Address: Evolutionary Ecology Unit, Department of Biology Lund University Lund SE‐22362 Sweden
3. Department of Biology and Mountain Lake Biological Station University of Virginia Charlottesville Virginia 22904
4. Department of Entomology University of Georgia Athens Georgia 30602
5. Department of Biological Sciences Virginia Tech Blacksburg Virginia 24060