1. Forest Research; Centre for Ecosystems, Society & Biosecurity; Northern Research Station Roslin Midlothian EH25 9SY UK
2. Biological & Environmental Sciences; University of Stirling; Stirling FK9 4LA UK
3. Plantlife Scotland; Balallan House Allan Park Stirling 2QG UK
4. Centre for Ecology and Hydrology; MacLean Bldg, Benson Ln Crowmarsh Gifford Wallingford Oxfordshire OX10 8BB UK