1. Key Laboratory of Fishery Drug Development of Ministry of Agriculture, Key Laboratory of Aquatic Animal Immune Technology of Guangdong Province, Pearl River Fisheries Research Institute Chinese Academy of Fishery Sciences Guangzhou China
2. College of Veterinary Medicine Northwest Agriculture and Forestry University Yangling China
3. College of Fisheries Tianjin Agriculture University Tianjin China
4. Institute of Infectology, Friedrich‐Loeffler‐Institut (FLI) Federal Research Institute for Animal Health Greifswald–Insel Riems Germany