1. SYSU‐Alberta Joint Lab for Biodiversity Conservation State Key Laboratory of Biocontrol and School of Life Sciences Sun Yat‐sen University Guangzhou 510275 China
2. Department of Ecology and Evolution University of California Los Angeles Los Angeles CA 90095 USA
3. Ministry of Education Key Laboratory of Western China's Environmental Systems College of Earth and Environmental Sciences Lanzhou University Lanzhou 730000 China
4. Department of Renewable Resources University of Alberta Edmonton AB T6G 2H1 Canada
5. Department of Plant and Environmental Sciences University of Copenhagen Frederiksberg DK‐1871 Denmark
6. CNRS ENFA; UMR5174 EDB (Laboratoire Evolution et Diversité Biologique) Université Paul Sabatier Toulouse France