1. The Key Laboratory of Plant Development and Environmental Adaptation Biology Ministry of Education College of Life Sciences Shandong University Qingdao Shandong266237 China
2. Shandong Provincial Key Laboratory of Plant Stress College of Life Sciences Shandong Normal University Jinan Shandong250014 China
3. The key Laboratory of Plant Stress Biology School of Life Science Henan University Kaifeng475004 China
4. Shandong Key Laboratory of Water Pollution Control and Resource Reuse School of Environmental Science and Engineering Shandong University Qingdao Shandong266237 China
5. Mendel Centre for Genomics and Proteomics of Plants Systems CEITEC MU – Central European Institute of Technology Masaryk University Brno004205 Czech Republic